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Coryphopteris Holtt.

Terrestrial ferns of small to moderate size. Rhizome an erect caudex, subarborescent (to 30 cm tall), rarely creeping, radially symmetric, dictyostelic, bearing many roots and broad, brown scales mostly without superficial and marginal acicular hairs, such scales also present on stipes. Fronds long-stipitate, the stipe dark throughout or paler distaly, with 2 vascular bundles at the base uniting upward into a single U-shaped vascular strand, lamina pinnate with pinnately deeply lobed pinnae, sometimes the basal lobes free, apex never pinna-like, the basal acroscopic lobe of pinna often enlarged and dentate, margins otherwise entire or toothed, basal pinnae little or not reduced, rarely gradually much-reduced, aerophores at bases of pinnae somewhat swollen, midribs of pinnae grooved above, but the grooves not confluent with rachis groove, reduced scales and unicellular, acicular hairs usually present on abaxial surfaces of axes, sometimes replaced by short, capitate hairs, sessile resinous glands (which may collapse on drying) +/- abundant in many species, adaxial surfaces with unicellular acicular hairs on axes sessile glands or hairs sometimes present between the veins, veins in a pinnate group in each pinna-lobe, free, simple, rarely once-forked, terminating in the margin above the sinus membrane. Sori round or asymmetric, indusiate, rarely exindusiate, indusium glandular or hairy; sporanjia on a short, 2-seriate stalk, lacking hairs or glands, sometimes with a sessile gland on the sporangium stalk, the annulus longitudinal interrupted. Spores monolete, pale, with a continuous wing and several (rarely many) cross wings.

Key modified ex Holttum 1981, F1. Males. ser. 2, 1:357 and 1977, Allertonia 1:196

Species in Papuasia

1a. Sessile glands and/or acicular hairs abundant between veins on upper surface of pinnae
Sessile glands and acicular hairs lacking from upper surface of pinnae
2a. Upper surface of lamina lacking acicular hairs between veins, glands and capitate hairs only
C. klossii
Upper surface of lamina bearing acicular hairs between veins, some glands or capitate hairs
C. obtusata
3a. Sessile glands present on lower surface generally, or at least on costules
Sessile glands lacking on lower surface except in c. fasciculata which may have a few glands on the costae
4a. Sori distinctly supramedial
C. diaphana
Sori medial to inframdial
5a. Basal half of lower pinnae bearing free or separately adnate pinnules
C. habbemensis
At most one free pinnule present at base of larges pinnae
6a. Sori mainly along one side of vein, rarely reniform
C. athyrioides
Sori mainly reniform, distal ones sometimes +/- athyrioid
7a. Lamina to 15 cm long; largest pinnae 3.0 x 0.8 cm
C. andreae

Lamina always much larger; pinnae in most cases to at least 5 x 1cm

8a. Lower pinnae, at least 2 pairs, reduced; basal pinnae not wider than the next above them
Basal pinnae wider and not shorter than those next above them
(C. pubirachis)


9a. Lamina to 25 cm long; pinnae 12-15 pairs (Pacific Islands)
L. pubirachis var. pubirachis
Lamina to 40 cm long; pinnae to c.20 pairs (Papuasia)
L. pubirachis var. major
10a. Six or more pairs of pinnae gradually reduced, lowest 1.5-2 cm long; stipe commonly to 15 cm long
C. squamipes
2-4 pairs of lower pinnae reduced, the lowest larger; stipe 20-25 cm long
11a. Indusia with many glands; fertile pinnae to 6 x 1.4cm; costules 3-3.5 cm apart; stipe scales to 0.5 mm wide
C. kolombangarae
Indusia with few hairs; fertile pinnae to 8.5 x 1.7cm; costule 4.5-55 mm apart; stipe scales to 1 mm wide
C. meiobasis
12a. Indusia lacking
C. engleriana
Indusia present
13a. Several pairs of free or separately adnate pinnules on lower pinnae
At most basal lobes free
14a. Lamina rigid; veins 4-5 pairs; indusia glabrous; no glands on costules and veins
C. stereohylla
Lamina soft; veins c.8 pairs; indusia with acicular hairs and glands; glands on costules and veins beneath
C. subbipinnata
15a. Upper surface bearing hairs between the veins
Upper surface between veins normally glabrous
16a. Stipe bearing copious hairs c.1 mm long
C. horizontalis
Stipe bearing hairs c.0.5 mm long in groove only
C. microlepigera
17a. Lower surface of rachis bearing acicular hairs, costae also
Lower surface of costae lacking (or almost lacking) acicular hairs
18a. Some hairs on upper surface of costae septate
C. lauterbachii
Hairs on upper surface of costae unicellular
19a. Several pairs of lower pinnae gradually reduced, longest 5.5cm, lowest 1-2cm
C. fasciculata
Lower pinnae not or little reduced
20a. Texture rigid; pinnae to 3 cm long; hairs on lower surface of rachis c.0.5 mm long
C. hubrechtensis
Textrue thin; pinnae to 6.5 cm long; hairs on lower surface of rachis c.0.2 mm long
C. brevipilosa
21a. Short capitate hairs present on lower surface of costae and costules and on indusia
C. oligolepia
Capitate hairs lacking or rare on lower surface or on indusia
22a. Sori supramedia; stipe scales 8-10 mm long
C. coriacea
Sori medial or inframedial; stipe scales not over 0.5 mm long
23a. Sori mostly athyrioid; pinnae to 2.5 mm long
C. ledermannii
Sori mostly not athyrioid; pinnae larger
24a. Caudex erect
Caudex slender, long-creeping
C. inopinnata
25a. Pinnae thin; no tangled hairs at base of stipe
Pinnae thick; tangled hairs often present at base of stipe
C. badia
26a. Pinnae caudate-acuminate
C. platyptera
Pinnae short-acuminate
C. subnigra