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Goniophlebium Bl.

Erect to pendulous epiphytes of moderate or large size. Rhizome long-creeping, bearing ovate, ciliate, apically attenuate, basally peltate, clathrate scales, dictyostelic, each vascular bundle surrounded by a sheath of sclerenchyma with additional strands of sclerenchyma in the cortex and medulla. Frond long-stipitate, the stipes articulate to phyllopodia, fibro-vascular bundles in a simple ring, the lamina pinnate, the pinnae stalked or subsessile, articulate to the rachis, shallowly or more obviously toothed, herbaceous, glabrous (finely hairy in juvenile plants), strongly costate with prominent veins, the main lateral veins close, bearing +/- excurrent lateral branches uniting to form 1 - 4 series of areoles decreasing in size towards the margin, each areole with a single (usually), simple, excurrent vein ending in a hydathode, the costal areoles with free veins arising from the base of the basiscopic main vein. Sporangia in discrete round sori in a single row either side of the costa, terminal on the included veinlets of the costal areoles, the receptacles immersed in the lamina and appearing as papillae on the upper surface of the lamina, or almost superficial, exindusiate, sometimes protected when young by peltate, dentate, clathrate paraphyses, seen as a ring around the sorus, annulus longitudinal, interrupted, of c.14 thickened cells. Spores monolete, smooth, translucent.

Species in Papuasia

1a, Large pendulous epiphytes; pinnae c.2 cm or more wide, bases narrowed or rounded, usually stalked 2
Medium size erect to arching epiphytes; pinnae c.1 cm wide, bases truncate or subcaudate, sessile or subsessile 3
2a. Dark, strongly toothed, clathrate, scale-like paraphyses present in an outer zone around each sorus G. percussum
Such paraphyses absent G. persicifolium
3a. Pinnae +/- auriculate at the base G. subauriculatum
Other ill-defined species such as: G. demersum