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(incl. Arcypteris)

Key ex Holttum 1974, Kew Bull. 29:341; 1951, Reinwardtia 1:193; 1951; Reinwardtia 1:176.

Species in Papuasia

1a. Fronds mostly bipinnatifid apart from basal pinnae, free pinnules, if present, few or attached by short-winged stalks to pinna-rachis; many additional areoles always present besides those along costae and costules (Arcypteris)


Fronds always amply bipinnate; additional areoles few or lacking (Pleocnemia s.s.)


2a. Fronds normally bipinnatifid, the largest sometimes bipinnate near the base; sori scattered irregularly, very abundant P. irregularis
Fronds, at least in the basal part, bipinnate with lobed pinnules; sori arranged in one row on each side of the costules of pinnule lobes 3
3a. Sori on all veins, +/- midway between the costule and margin, usually distinct P. macrodonta
Sori close to the margin, lacking from basal veins often +/- confluent P. andaiensis
4a. Sori indusiate 5
Sori exindusiate 7
5a. Pinnules of middle pinnae lobed c.3/5 towards costule; lobes distinctly falcate with small teeth at ends of veins; sinuses narrow, sinus teeth small but elongate P. conjugata
Pinnules of middle pinnae usually lobed more deeply; sinuses wider, sinus teeth +/- triangular 6
6a. Pinnules hardly over 2 cm wide, their lobes crenate P. seranensis
Pinnules commonly 2.5-3 cm wide, their lobes lobed, veins in lobules twice-forked P. cumingiana
7a. Pinnules lobed c.2/3 towards costa; lobes entire or at most minutely toothed P. leuzeana
Pinnules lobed more deeply or not; lobes crenate 8
8a. Pinnules of middle pinnae c. 2 cm wide, lobed 1/2-2/3, lobes hardly falcate, blunt 9
Pinnules of middle pinnae commonly 3.5-4.5 cm wide, lobed 1-2 mm from costa, lobes distinctly falcate, acute P. megaphylla
9a. Pinnules lobes c.1/2 way to costa P. dimidilobata
Pinnules pinnate at base (at least lower ones) P. tripinnata

Note: Holttum (1974) considers that P. dimidilobata includes P. tripinnata