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Key ex Holttum 1985, in litt.

Species in Papuasia

1a. Veins all free T. dissecta
Veins anastomosing 2
2a. Veins forming regular costa areoles lacking free included veinlets; other areoles, if present, with few short, unbranched veinlets 3
Veins copiously anastomosing with many, often branched, free veinlets in areoles 11
3a. Upper surface bearing hairs between the veins 4
Upper surface lacking hairs between veins, except near margins 7
4a. Indusia absent of very small 5
Indusia conspicuous 6
5a. Stipe and rachis dark, glossy; axes or frond scaly on abaxial surface, not hairy "T. squamipes"
Stipe and rachis not dark and glossy; axes of frond +/- hairy on abaxial surface, not scaly T. ferruginea
6a. Additional areoles ocmmonly present below sinuses between lobes of pinnae or pinnules; hairs between veins on upper surface very thick T. pubescens
Additional areoles below sinuses lacking or rare; hairs on upper surface slender; limestone plants T. devexa
7a. Suprabasal pinnae with several pairs of free or adnate pinnules; sterile and fertile fronds isomorphous 8
Suprabasal pinnae lacking free pinnules; fronds +/- dimorphous 10
8a. Rachises dark and glossy, bearing numerous narrow scales on abaxial surfaces "T. nabirensis"
Rachises lacking scales on aboxial surfaces 9
9a. Basal acroscopic lobes of pinnules little longer than basal basiscopic ones "T. nesiotica"
Basal acroscopic lobes of pinnules much longer than basal basiscopic ones T. kingii
10a. Fronds opaque, smaller veins not visible on surfaces; buds present at bases of pinnae T. bamleriana
Fronds not opaque; veins all distinct; no buds. T. brachiata
11a. Lamina simple, its base cordate T. novoguineensis
Lamina deeply 3 - 5-lobed or pinnate 12
12a. Lamina deeply 3 - 5-lobed, its base cordate 13
Lamina lobed to a winged rachis; stipe also winged 15
12c. Lamina consisting of free pinnae, at least at its base; stipe unwinged 17
13a. Sori indusiate, near margins; both surfaces copiously short-hairy T. craspedocarpa
Sori exindusiate, irregularly scattered, sometimes coalescent 14
14a. Lower surface of costae glabrous; rather many coalescent sori; basal scales short T. cristovalensis
Lower surface of costae hairy; basal scales 15 mm or more long; fewer coalescent sori "T. croftii" (juvenille)
15a. Sori in one row each side of main veins, +/- prominent on the upper surface; indusia large T. decurrens
Sori exindusiate, not thus arranged 16
16a. Sori rather regularly arranged in rows parallel to cross veins which join main veins; wing commonly 10-15 mm wide on each side of the rachis T. beccariana
Sori very small, irregularly scattered; wing of rachis much narrower T. schultzei
17a. Suprabasal pinnae entire or subentire 18
Suprabasal pinnae deeply lobed 30
18a. Sori in one row on each side of main veins 19
Sori not in one row on each side of main veins 26
19a. Indusium large and conspicuous; sori uniform 20
Indusium small and thin or lacking; sori sometimes +/- enlongate or coalescing 25
20a. At least basal pinnae auricled on their acroscopic side T. barclayi
No pinnae auricled on the acroscopic side 21
21a. Stipe and rachis dark and glossy 22
Stipe and rachis not dark and gloosy 23
22a. Bases of suprabasal pinnae broadly rounded; scales at base of stipe rigid, erect, dark twisted "T. exauriculata"
Bases of suprabasal pinnae cuneate; scales at base of stipe otherwise "T. gymorachis"
23a. Basal pinnae forked; indusia peltate or round-renform, receptacle not elongate along the vein 24
Basal pinnae not forked; indusia +/- ellipsoid, receptacle elongate along the vein T. pleiosora
24a. Apical lamina at most trilobed; lower surface of costae glabrous; indusia very firm, round reniform "T. isomorpha"
Apical lamina commonly multilobed; lower surface of costae minutely hairy; indusium thinner, peltate to reniform T. crenata
25a. Sori sometimes elongate or coalescent parallel to the main veins; fronds not dimorphous; pinnae to c.2 cm wide "T. refescens"
Sori (especially the basal ones) elongate and sometimes coalescent at right angles to main veins; fronds usually dimorphous T. siifolia
26a. Apical lamina trilobed, with 1 or more buds on the upper surface of its midrib; sori small, irregularly arranged, mostly on small veinlets in areoles T. angulata
Apical lamina lacking buds; sori mostly not on free veinlets in areoles 27
27a. Caudex creeping, to 6 mm diameter, basal pinnae unlobed, to c. 25 x 3cm T. menyanthidis
Caudex erect or ascending, with tufted fronds; basal pinnae shorter, usually lobed 28
28a. Fertile pinnae c.7 mm wide, much narrower than sterile, with simpler venation; sori nearly all coalescent, some coenosori linear and submarginal T. teratocarpa
Fertile pinnae conform with sterile 29
29a. Basal pinnae 4-5 cm wide above basal lobe; buds present at their bases "T. croftii"
Basal pinnae to 2 cm wide above basal lobe; no buds at their bases "T. rheophytica"
30a. Stipe and rachis dark 31
Stipe and rachis not dark T. brachiata
31a. Apical lamina many-lobed 32
Apical lamina trilobed T. barclayi
32a. Fronds dimorphous; areoles containing sori filling whole area of lobes of fertile pinnae T. durvillei
Fronds not dimorphous; many areoles not containing sori; limestone plants T. melanorachis


Monotypic genus...S. aurita

Hemigramma or gen. Nov. aff.

Key ex Copeland 1928, Philip. J. Sci. 37:403.

Species in Papuasia

1a. Simple, or pinnate with rachis mostly winged H. hollrungii
Pinnate, rachis mostly wingless H. grandifolia